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Cu + is taken up by a high-affinity transporter on the hepatocyte cell membrane, incorporated into ceruloplasmin, released into the bloodstream, delivered to all organs, and eventually excreted via the biliary route [14]

doi: 10.1016/j.spen.2020.100835

In its neutral form, the imidazole side chain has an "ambidextrous" nature, since the nitrogen without a hydrogen is nucleophilic and can act as a hydrogen bond acceptor, while the nitrogen with the hydrogen bond is electrophilic and can act as a H-bond donor

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Lin Y-K, Yang S-H, Chen C-C, Kao H-C, Fang J-Y

"Cobalamin Deficiency: Clinical Picture and Radiological Findings"